Geniostoma hymenopodum

(O.Deg. & Sherff) Byng & Christenh. (2018)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Asterids Order: Gentianales Family: Loganiaceae Genus: Geniostoma

kāmakahala [kamakahala]

Description

Key Characters:

Growth Form: Branched shrubs 1–3 m tall.

Stems: Stems obscurely angled, moderately short–hispid.

Roots:

Leaves: Leaves simple. Opposite. Blades oblanceolate to elliptic or sometimes obovate, 4–15 cm long, (1.5–)2–4.5 cm wide. Apex acute to bluntly short-acuminate. Base gradually or strongly narrowed to a subtruncate base or base sometimes broadly cuneate. Upper surfaces glabrous; lower surfaces glabrous or sparsely short–hispid, sometimes only along veins; leaves yellowish green to medium green on upper surface; lower surfaces pale green, coriaceous. Margins ± slightly revolute. Upper surface with veins slightly impressed. Petioles 0.2–1 cm long, glabrous. Stipules interpetiolar, may also be intrapetiolar, membranous, connate into a cup (3–)5–14 mm long, adnate in lower ⅓ to petioles, quickly splitting, glabrous or short–hispidulous near base, caducous.

Flowers: Flowers in inflorescences terminal, flowers 3–12 in sessile cymes, pedicels 1–3 mm long, glabrous or sparsely short–hispidulous, bracts linear–spatulate to linear–filiform, 4–11 mm long, glabrous, bracteoles filiform, 3–5 mm long, glabrous. Flowers bisexual (perfect) or functionally unisexual, actinomorphic. Calyx 5-merous, sepals united at base, calyx lobes imbricate, distinctly unequal, spatulate to oblanceolate, the 3 outer ones (4–)6–12 mm long, (0.8–)2–7 mm wide, the 2 inner ones (4–)5–10 mm long, 0.6–4 mm wide, inconspicuously 3–7–nerved, glabrous or sparsely short–hispid, inner surface of calyx often with colleters at the base, margin mostly fimbriate. Corolla 5-merous, campanulate to rotate, pale orange, 9.5–16 mm long, the tube slightly constricted near throat, 6.3–11 mm long, glabrous or sometimes sparsely short–hispidulous, the lobes narrowly deltate, 3.2–6 mm long, apex acute, white short–hirsute on the tube within and on lower part of lobes, lobes imbricate or contorted in bud. Stamens 5, alternipetalous, attached to mouth of corolla, exserted, usually recurved after anthesis, glabrous or hairy; filaments mostly short; anthers 2-celled, introrse, connective often extended to form an apical appendage, opening by longitudinal slits. Ovary superior or partly inferior, carpels hairy or glabrous, 2– or 3-locular, sometimes partitions imperfect in upper part of ovary, ca. 8–10 mm long, glabrous; ovules numerous, amphitropous, placentation axile; style terminal, mostly short; stigma clavate, ellipsoid, to globular, sometimes ± oblong, about as large as ovary.

Fruit: Capsules ovoid–trigonous; 12–14 mm long; 3–valved; septicidal to sub–septifragal; valves keeled in upper part; apex with a beak ca. 3 mm long; maturing to green; with or without a black–purple tinge; usually becoming brown–black when over–mature. Seeds not seen.

Ploidy:

Habitat: Occurring in wet forest.

Elevation Range: 600–860 m.

Historical Distribution

Uses and Culture

USES

Natural History

Statewide Status

Endemic

Island Status

O'ahu Endemic

Dispersal Agents


Pollinators

Specimens

Bibliography

Name Published In: Global Fl. 4: 145 (2018)

Other References

Wagner et al. 1990:860 (O [as Labordia sessilis]); Gibbons et al. 2012/Christenhusz et al. 2018:145 (Syn. L. sessilis = G. hymenopodum)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Geniostoma hymenopodum Koolau Range; Puu Kaaumakua Waikane-Waipio Mossy crest Preserved_Specimen St.John, H. 18000 Oahu BISH 1937-05-30
2 Geniostoma hymenopodum Poamoho Solitary, in dense uluhe thicket in native forest. Preserved_Specimen Takeuchi, W.N. 2227 Oahu BISH 1985-06-09
3 Geniostoma hymenopodum Kawailoa C.C.C. Trail Rain-swept, shrubby summit. Preserved_Specimen Degener, O. 12059 Oahu BISH 1938-07-03
4 Geniostoma hymenopodum Koolau Mountains; Waipio. S ridge, Kipapa Gulch Wet denuded ridge. Preserved_Specimen Hosaka, E.Y. 1139 Oahu BISH 1933-08-06
5 Geniostoma hymenopodum Aiea C.C.C. Trail Wind & rain-swept summit. Preserved_Specimen Degener, O. 11550 Oahu BISH 1937-12-06
6 Geniostoma hymenopodum Main divide of Koolau Mountains; Heeia Top of cliffs at head of Haiku Valley Wet, bushy, wind-swept ridge. Preserved_Specimen Fosberg, F.R. 13860 Oahu BISH 1937-05-23
7 Geniostoma hymenopodum Koolau Mountains; Kahana Waikane-Schofield Trail Wet, steep, exposed bushy slope. Preserved_Specimen Fosberg, F.R. 12249 Oahu BISH 1935-07-27
8 Geniostoma hymenopodum Koolau range; Waikane-Schofield Trail In rain forest. Preserved_Specimen Cowan, R.S. 12 Oahu BISH 1946-07-29
9 Geniostoma hymenopodum Honolulu District. Pia gulch, top of Hawaiiloa trail Linearis wet forest. With Cheirodendron trigynum, Broussaisia arguta, Diplazium sandwichianum, Mysine lessertiana, Cibotium, Antidesma platyphylla, Pritchardia martiik, Syzgium sandwicensis, Bobea elatior, Pipturus, Scaev Preserved_Specimen Perlman, S. 23537 Oahu BISH 2013-06-18
10 Geniostoma hymenopodum Koolau Range; Puu Kainapuaa Occasional, on wooded slope. Preserved_Specimen Hosaka, E.Y. 1918 Oahu BISH 1937-05-31
11 Geniostoma hymenopodum Opposite Wiliwilinui Ridge, overlooking Waimanalo Fog-swept, precipitous slope Preserved_Specimen Degener, O. 19709 Oahu BISH 1949-06-27
12 Geniostoma hymenopodum Koolau Range. Divide between Waikane & Waikakalaua Gulches On wooded slope below ridge. Preserved_Specimen Hosaka, E.Y. 1880 Oahu BISH 1937-05-30
13 Geniostoma hymenopodum Poamoho In open clearing near the summit. Preserved_Specimen Takeuchi, W.N. 2228 Oahu BISH 1985-06-09
14 Geniostoma hymenopodum Puu Kahuauli In open forest. Preserved_Specimen Degener, O. 19659 Oahu BISH 1949-06-19
15 Geniostoma hymenopodum Kaluanui Valley By stream Preserved_Specimen Hume, E.P. 556 Oahu BISH 1932-05-08
16 Geniostoma hymenopodum Koolau Mountains crest; main divide. Between Waianu Gulch & Kipapa Gulch Wet forest. Preserved_Specimen Fosberg, F.R. 13914 Oahu BISH 1937-05-30
17 Geniostoma hymenopodum Waipio; Kipapa Gulch Wooded ridge. Preserved_Specimen Fosberg, F.R. 9723 Oahu BISH 1933-08-06
18 Geniostoma hymenopodum Koolau Mountains; Kipapa Gulch On denuded ridge. Preserved_Specimen Hosaka, E.Y. 1139 Oahu BISH 1933-08-06
19 Geniostoma hymenopodum Poamoho In open clearing near summit Preserved_Specimen Takeuchi, W.N. 2228 Oahu BISH 1985-06-09
20 Geniostoma hymenopodum Waipio; Kipapa-Waiawa Ridge Metrosideros forest. Preserved_Specimen Grant, M.L. 7199 Oahu BISH 1934-08-07
21 Geniostoma hymenopodum Koolau range; Waikane-Schofield Trail In rain forest. Preserved_Specimen Cowan, R.S. 12 Oahu BISH 1946-07-29
22 Geniostoma hymenopodum N. Koolau Mountains; Punaluu (Castle) Trail. Along upper Kalaunui Stream In wet Metrosideros forest with Euphorbia rockiana, Scaevola mollis, Cheirodendron platyphyllum, Broussaisia, Pipturus. Preserved_Specimen Carlquist, S. 1737 Oahu BISH 1965-07-06
23 Geniostoma hymenopodum Waipio; Kipapa Gulch Exposed ridge. Preserved_Specimen Fosberg, F.R. 9741 Oahu BISH 1933-08-06
24 Geniostoma hymenopodum Poamoho Summit divide near Poamoho trail terminus In windbeaten habitat. Preserved_Specimen Takeuchi, W.N. 2349 Oahu BISH 1985-06-09
25 Geniostoma hymenopodum Koolau Mountains crest; main divide. Between Waianu Gulch & Kipapa Gulch Wet forest. Preserved_Specimen Fosberg, F.R. 13917 Oahu BISH 1937-05-30
26 Geniostoma hymenopodum Poamoho Trail At wind-swept open top. Preserved_Specimen Degener, O. 27387 Oahu BISH 1961-04-30
27 Geniostoma hymenopodum Waipio; Kipapa Gulch Exposed ridge. Preserved_Specimen Fosberg, F.R. 9725 Oahu BISH 1933-08-06
28 Geniostoma hymenopodum Koolau Mountains; Moanalua Valley; Summit crest ridge In Metrosideros-Dicranopteris linearis lowland wet forest with Antidesma, Clermontia, Cyrtandra, Zanthoxylum, Myrsine, Melicope, Pritchardia, Psychotria, Tetraplasandra, Cheirodendron, Bobea. Preserved_Specimen Perlman, S. 5934 Oahu BISH 1987-06-07
29 Geniostoma hymenopodum Poamoho Solitary, in dense uluhe thicket in native forest. Preserved_Specimen Takeuchi, W.N. 2227 Oahu BISH 1985-06-09
30 Geniostoma hymenopodum Waimanalo Preserved_Specimen Degener, O. 19709 Oahu BISH 1949-06-27
31 Geniostoma hymenopodum Koolau Range, near summit of Kipapa Trail Preserved_Specimen Kerr, M. 21662 Oahu BISH 1947-06-21
32 Geniostoma hymenopodum Summit trail between Laie & Castle Trails Preserved_Specimen Kerr, M. 2901 Oahu BISH 1946-06-02
33 Geniostoma hymenopodum Punaluu Along Pig-God trail Preserved_Specimen Degener, O. 10335 Oahu BISH 1932-06-22
34 Geniostoma hymenopodum Punaluu. Near top of range opposite marsh trail Preserved_Specimen Russ, G.W. Oahu BISH 1933-07-24
35 Geniostoma hymenopodum Nuuanu Lanihuli Preserved_Specimen Russ, G.W. Oahu BISH 1933-07-19
36 Geniostoma hymenopodum Kipapa-Waiahole, summit Preserved_Specimen Kerr, M. Oahu BISH 1946-07-04